Díaz-Tapia, P.; McIvor, L.; Freshwater, D. W.; Verbruggen, H.; Wynne, M. J.; Maggs, C. A. (2017). The genera Melanothamnus Bornet & Falkenberg and Vertebrata S.F. Gray constitute well-defined clades of the red algal tribe Polysiphonieae (Rhodomelaceae, Ceramiales).
European Journal of Phycology. 52(1): 1-30.