Lönnberg, E. (1906). Contributions to the fauna of South Georgia. I. Taxonomic and biological notes on vertebrates.
Kongliga Vetenskaps-Academiens Handlingar, Stockholm n. s. Series 4, v. 40 (no. 5): 1-104, Pls. 1-12. [Fishes on pp. 91-100, no plates.].