Copepoda name details
Epactophanes richardi Mrázek, 1893
238113 (urn:lsid:marinespecies.org:taxname:238113)
alternative representation
Species
Canthocamptus papuanus Daday, 1901 · unaccepted > junior subjective synonym (Synonym of Epactophanes richardi...)
Synonym of Epactophanes richardi after Lang (1948) and Defaye & Dussart (2011)
Epactophanes muscicola (Richters, 1900) · unaccepted
Epactophanes quadrispinosus (Richters, 1908) · unaccepted
Epactophanes richardi muscicola (Richters, 1900) · unaccepted
Epactophanes richardi typica Kiefer, 1925 · alternative representation
Epactophanes wolfi (Richters, 1907) · unaccepted
Moraria muscicola Richters, 1900 · unaccepted (synonym)
Moraria quadrispinosa Richters, 1908 · unaccepted (synonym)
- Subspecies Epactophanes richardi aculeatus Lang, 1934
- Subspecies Epactophanes richardi arcticus Lang, 1934
- Subspecies Epactophanes richardi bidens Lang, 1934
- Subspecies Epactophanes richardi intermedius Borutzky, 1925
- Subspecies Epactophanes richardi menzeli Chappuis, 1931
- Subspecies Epactophanes richardi richardi Mrázek, 1893
- Subspecies Epactophanes richardi tuberculatus Lastochkin, 1924
- Subspecies Epactophanes richardi antarcticus (Richters, 1907) accepted as Epactophanes antarcticus (Richters, 1907)
- Subspecies Epactophanes richardi muscicola (Richters, 1900) accepted as Epactophanes richardi Mrázek, 1893 represented as Epactophanes richardi richardi Mrázek, 1893
- Subspecies Epactophanes richardi typica Kiefer, 1925 represented as Epactophanes richardi Mrázek, 1893 represented as Epactophanes richardi richardi Mrázek, 1893
Mrázek, A. (1893). Beitrag zur Kenntnis der Harpacticidenfauna des Süsswassers. [Contribution to the knowledge of the freshwater harpacticid fauna.]. <em>Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere.</em> 7(1): 89-130, pls. 4-7. (volume date published in 1894 however the first few issues were published in 20-v-1893)., available online at https://www.biodiversitylibrary.org/page/39199167
page(s): 108 [details]
page(s): 108 [details]
Walter, T.C.; Boxshall, G. (2025). World of Copepods Database. Epactophanes richardi Mrázek, 1893. Accessed at: https://www.marinespecies.org/copepoda/aphia.php?p=taxdetails&id=238113 on 2025-07-01
Date
action
by
original description
Mrázek, A. (1893). Beitrag zur Kenntnis der Harpacticidenfauna des Süsswassers. [Contribution to the knowledge of the freshwater harpacticid fauna.]. <em>Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere.</em> 7(1): 89-130, pls. 4-7. (volume date published in 1894 however the first few issues were published in 20-v-1893)., available online at https://www.biodiversitylibrary.org/page/39199167
page(s): 108 [details]
basis of record Fiers, F. & V. Ghenne. (2000). Cryptozoic copepods from Belgium: diversity and biogeographic implications. <em>Belgian Journal of Zoology.</em> 103(1): 11-19. [details] Available for editors
[request]
additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors [request]
additional source Brehm, V. (1912). Die Entomostraken der Danmark-Expedition. [The Entomostraca of the Denmark Expedition.]. <em>Meddelelser om Gronland, In: Danmark-Expeditionen til Gronlands Nordostkyst 1906-1908, Kobenhavn.</em> 5(5)45:305-317, pls.18-19. (1911). [details]
additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors [request]
additional source Dussart, B.H. (1967). Les Copépodes des eaux continentales d'Europe occidentale. Tome I: Calanoïdes et Harpacticoïdes. [Copepods of the continental waters of Western Europe. Volume I: Calanoids and Harpacticoids.]. <em>N. Boubée et Cie, Paris.</em> 500 pp.
page(s): 290 [details] Available for editors [request]
additional source Franke, U. (1989). Katalog zur Sammlung limnischer Copepoden von Prof. Dr. Friedrich Kiefer (Mit einem Gesamtverzeichnis seiner Publikationen). [Catolog of the freshwater copepods of Dr. Friedrich Kiefer.]. <em>Carolinea Beiheft, Karlsrühe.</em> 5:1-433, figs. 1-2.
note: Catalog vial number(s): 3532, 77 [details] Available for editors [request]
additional source Ishida, T. (1982). Further notes on the freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 36:49-61, figs. 1-9. (iii-1982, Japanese with English summary.) [details] Available for editors [request]
additional source Ishida, T. (1987). Freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 41:77-119. (Japanese summary.) [details] Available for editors [request]
additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors [request]
additional source Ishida, T. (1998). Freshwater copepods from the east coast of the Kamchatka Peninsula, Russia. In: Dahms, H.U., T. Glatzel, H.J. Hirche, S. Schiel & H.K. Schminke, (eds.). Proceedings of the 6th International Conference on Copepoda, Journal of Marine Systems, Spec. Vol. 15(1-4):391-396. [details] Available for editors [request]
additional source Menzel, R. (1914). Zur Kenntnis von Moraria muscicola Richters nebst Hinweis auf deren Verwandtschaft mit Epactophanes richardi Mrázek. [On the knowledge of Moraria muscicola Richters including reference to its relationship with Epactophanes richardi Mrázek.]. <em>Zoologischer Anzeiger.</em> 43(13):615-620, figs. 1-5. (17-iii-1914). [details]
additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors [request]
additional source Kiefer, F. (1924). 3. Beiträge zur Copepodenkunde. 1. Canthocamptus pygmaeus Sars. 2. Ein neuer Fundort von Diaptomus denticornis Wierz. im Schwartzwald. 3. Epactophanes richardi Mràzek. <em>Zoologischer Anzeiger.</em> 59(7-8):200-204, figs. 1-8. (5-v-1924). [details] Available for editors [request]
additional source Kiefer, F. (1960). Psammobionte Ruderfusskrebse (Crust. Cop.) aus dem Gebiet der Unterweser und von der Insel Helgoland. [Psammobiont copepods (Crust. Cop.) from the Lower Weser region and from the island of Heligoland. <em>Zoologischer Anzeiger.</em> 165(1-2):30-36, figs. 1-39. (vii-1960). [details] Available for editors [request]
additional source Kulhavy, V. (1957). Plazivky (Copepoda-Harpacticoidea) jihoceskych mechu. [Harpacticoid copepods from South Bohemian mosses.]. <em>Acta Musei Nationalis, Prahba, [ Casopis Národniho Musea].</em> 126(1):35-43, figs. 1-6. (Czech with English summary.).
page(s): 41 [details] Available for editors [request]
additional source Michailova-Neikova, M. (1963). Epactophanes richardi Mrázek. <em>Priroda, Moscow.</em> 12(4):101-102. [details]
additional source Monchenko, V.I. (1964). Novi dlya Ukraini veslonogi rakopodibni (Copepoda): Parastenocaris brevipes ta Epactophanes richardi. [New copepods for the Ukraine: Parastenocaris brevipes and Epactophanes richardi.]. <em>Dopovidi Akademiyi Nauk Ukrayins'koiy RSR, Kiev.</em> 1964(4):545-549. (Ukrainian with Russian and English summaries). [details] Available for editors [request]
additional source Okuneva, G.L. (1989). Garpaktitsidy ozera Bakala. [The harpacticoids of Lake Baikal.]. <em>Izdatel'stvo Instituta, Irkutsk.</em> 150 pp. (In Russian.). [details] Available for editors [request]
additional source Lewis, M.H. (1980). Systematic studies on the New Zealand freshwater Harpacticoida (Crustacea: Copepoda). <em>Ph.D. Thesis, University of Auckland, Auckland, New Zealand.</em> 314 pp., available online at https://hdl.handle.net/2292/1599 [details]
additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors [request]
additional source Defaye, D. & B.H. Dussart. (2011). World Directory of Crustacea Copepoda III - Harpacticoida IV - Gelyelloida. <em>Backhuys Publishers, Margraf Publishers GmbH.</em> 450 pp.
note: "The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); ...
[details] Available for editors [request]
additional source Kiefer, F. (1963). Über einige Copepoda Harpacticoida aus grabungen an norditalienischen Fliessgewässern. [On some Copepoda Harpacticoida from excavations at northern Italian rivers.]. <em>Estratto dalle Memorie del Museo Civico di Storia Naturale, Verona.</em> 11:1-10. (15 Jul 1963). [details] Available for editors [request]
additional source Marin, I., D. Sharafutdinova, T. Mukhina, A. Chernyshev & A. Novikov. (2022). The record of Epactophanes richardi Mrázek, 1893 (Harpacticoida: Canthocamptidae) in soil of the Russian Far-Eastern forest. <em>Journal of Asia-Pacific Biodiversity.</em> 15:454-458., available online at https://doi.org/10.1016/j.japb.2022.05.002 [details] Available for editors [request]
page(s): 108 [details]
basis of record Fiers, F. & V. Ghenne. (2000). Cryptozoic copepods from Belgium: diversity and biogeographic implications. <em>Belgian Journal of Zoology.</em> 103(1): 11-19. [details] Available for editors
[request]additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors
[request]additional source Brehm, V. (1912). Die Entomostraken der Danmark-Expedition. [The Entomostraca of the Denmark Expedition.]. <em>Meddelelser om Gronland, In: Danmark-Expeditionen til Gronlands Nordostkyst 1906-1908, Kobenhavn.</em> 5(5)45:305-317, pls.18-19. (1911). [details]
additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors
[request]additional source Dussart, B.H. (1967). Les Copépodes des eaux continentales d'Europe occidentale. Tome I: Calanoïdes et Harpacticoïdes. [Copepods of the continental waters of Western Europe. Volume I: Calanoids and Harpacticoids.]. <em>N. Boubée et Cie, Paris.</em> 500 pp.
page(s): 290 [details] Available for editors
[request]additional source Franke, U. (1989). Katalog zur Sammlung limnischer Copepoden von Prof. Dr. Friedrich Kiefer (Mit einem Gesamtverzeichnis seiner Publikationen). [Catolog of the freshwater copepods of Dr. Friedrich Kiefer.]. <em>Carolinea Beiheft, Karlsrühe.</em> 5:1-433, figs. 1-2.
note: Catalog vial number(s): 3532, 77 [details] Available for editors
[request]additional source Ishida, T. (1982). Further notes on the freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 36:49-61, figs. 1-9. (iii-1982, Japanese with English summary.) [details] Available for editors
[request]additional source Ishida, T. (1987). Freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 41:77-119. (Japanese summary.) [details] Available for editors
[request]additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors
[request]additional source Ishida, T. (1998). Freshwater copepods from the east coast of the Kamchatka Peninsula, Russia. In: Dahms, H.U., T. Glatzel, H.J. Hirche, S. Schiel & H.K. Schminke, (eds.). Proceedings of the 6th International Conference on Copepoda, Journal of Marine Systems, Spec. Vol. 15(1-4):391-396. [details] Available for editors
[request]additional source Menzel, R. (1914). Zur Kenntnis von Moraria muscicola Richters nebst Hinweis auf deren Verwandtschaft mit Epactophanes richardi Mrázek. [On the knowledge of Moraria muscicola Richters including reference to its relationship with Epactophanes richardi Mrázek.]. <em>Zoologischer Anzeiger.</em> 43(13):615-620, figs. 1-5. (17-iii-1914). [details]
additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors
[request]additional source Kiefer, F. (1924). 3. Beiträge zur Copepodenkunde. 1. Canthocamptus pygmaeus Sars. 2. Ein neuer Fundort von Diaptomus denticornis Wierz. im Schwartzwald. 3. Epactophanes richardi Mràzek. <em>Zoologischer Anzeiger.</em> 59(7-8):200-204, figs. 1-8. (5-v-1924). [details] Available for editors
[request]additional source Kiefer, F. (1960). Psammobionte Ruderfusskrebse (Crust. Cop.) aus dem Gebiet der Unterweser und von der Insel Helgoland. [Psammobiont copepods (Crust. Cop.) from the Lower Weser region and from the island of Heligoland. <em>Zoologischer Anzeiger.</em> 165(1-2):30-36, figs. 1-39. (vii-1960). [details] Available for editors
[request]additional source Kulhavy, V. (1957). Plazivky (Copepoda-Harpacticoidea) jihoceskych mechu. [Harpacticoid copepods from South Bohemian mosses.]. <em>Acta Musei Nationalis, Prahba, [ Casopis Národniho Musea].</em> 126(1):35-43, figs. 1-6. (Czech with English summary.).
page(s): 41 [details] Available for editors
[request]additional source Michailova-Neikova, M. (1963). Epactophanes richardi Mrázek. <em>Priroda, Moscow.</em> 12(4):101-102. [details]
additional source Monchenko, V.I. (1964). Novi dlya Ukraini veslonogi rakopodibni (Copepoda): Parastenocaris brevipes ta Epactophanes richardi. [New copepods for the Ukraine: Parastenocaris brevipes and Epactophanes richardi.]. <em>Dopovidi Akademiyi Nauk Ukrayins'koiy RSR, Kiev.</em> 1964(4):545-549. (Ukrainian with Russian and English summaries). [details] Available for editors
[request]additional source Okuneva, G.L. (1989). Garpaktitsidy ozera Bakala. [The harpacticoids of Lake Baikal.]. <em>Izdatel'stvo Instituta, Irkutsk.</em> 150 pp. (In Russian.). [details] Available for editors
[request]additional source Lewis, M.H. (1980). Systematic studies on the New Zealand freshwater Harpacticoida (Crustacea: Copepoda). <em>Ph.D. Thesis, University of Auckland, Auckland, New Zealand.</em> 314 pp., available online at https://hdl.handle.net/2292/1599 [details]
additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors
[request]additional source Defaye, D. & B.H. Dussart. (2011). World Directory of Crustacea Copepoda III - Harpacticoida IV - Gelyelloida. <em>Backhuys Publishers, Margraf Publishers GmbH.</em> 450 pp.
note: "The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); ...
"The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); Damian-Georgescu (1970) and Tai & Song (1979) considered them as synonyms of the nominal species which appears to have an important morphological variability. Borutzky (1952) did not agree with this opinion" (Defaye & Dussart, 2011: 115).
[request]additional source Kiefer, F. (1963). Über einige Copepoda Harpacticoida aus grabungen an norditalienischen Fliessgewässern. [On some Copepoda Harpacticoida from excavations at northern Italian rivers.]. <em>Estratto dalle Memorie del Museo Civico di Storia Naturale, Verona.</em> 11:1-10. (15 Jul 1963). [details] Available for editors
[request]additional source Marin, I., D. Sharafutdinova, T. Mukhina, A. Chernyshev & A. Novikov. (2022). The record of Epactophanes richardi Mrázek, 1893 (Harpacticoida: Canthocamptidae) in soil of the Russian Far-Eastern forest. <em>Journal of Asia-Pacific Biodiversity.</em> 15:454-458., available online at https://doi.org/10.1016/j.japb.2022.05.002 [details] Available for editors
[request]
Present Inaccurate Introduced: alien Containing type locality
Nontype NHMW 22928, geounit Malaysia [details]
To Barcode of Life (17 barcodes)
To Biodiversity Heritage Library (1 publication) (from synonym Canthocamptus papuanus Daday, 1901)
To Biodiversity Heritage Library (17 publications) (from synonym Moraria muscicola Richters, 1900)
To Biodiversity Heritage Library (2 publications) (from synonym Moraria quadrispinosa Richters, 1908)
To Biodiversity Heritage Library (24 publications)
To Biodiversity Heritage Library (3 publications) (from synonym Epactophanes muscicola (Richters, 1900))
To European Nucleotide Archive, ENA (Epactophanes richardi)
To GenBank (23 nucleotides; 14 proteins)
To PESI
To USNM Invertebrate Zoology Arthropoda Collection (13 records)
To ITIS
To Biodiversity Heritage Library (1 publication) (from synonym Canthocamptus papuanus Daday, 1901)
To Biodiversity Heritage Library (17 publications) (from synonym Moraria muscicola Richters, 1900)
To Biodiversity Heritage Library (2 publications) (from synonym Moraria quadrispinosa Richters, 1908)
To Biodiversity Heritage Library (24 publications)
To Biodiversity Heritage Library (3 publications) (from synonym Epactophanes muscicola (Richters, 1900))
To European Nucleotide Archive, ENA (Epactophanes richardi)
To GenBank (23 nucleotides; 14 proteins)
To PESI
To USNM Invertebrate Zoology Arthropoda Collection (13 records)
To ITIS
