RAS name details
Epactophanes richardi Mrázek, 1893
238113 (urn:lsid:marinespecies.org:taxname:238113)
alternative representation
Species
- Subspecies Epactophanes richardi richardi Mrázek, 1893
- Subspecies Epactophanes richardi antarcticus (Richters, 1907) accepted as Epactophanes antarcticus (Richters, 1907)
Mrázek, A. (1893). Beitrag zur Kenntnis der Harpacticidenfauna des Süsswassers. [Contribution to the knowledge of the freshwater harpacticid fauna.]. <em>Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere.</em> 7(1): 89-130, pls. 4-7. (volume date published in 1894 however the first few issues were published in 20-v-1893)., available online at https://www.biodiversitylibrary.org/page/39199167
page(s): 108 [details]
page(s): 108 [details]
Walter, T.C.; Boxshall, G. (2025). World of Copepods Database. Epactophanes richardi Mrázek, 1893. Accessed through: RAS (Eds.) (2025) Register of Antarctic Species at: https://ras.biodiversity.aq/aphia.php?p=taxdetails&id=238113 on 2026-09-09
RAS (Eds.) (2026). Register of Antarctic Species. Epactophanes richardi Mrázek, 1893. Accessed at: https://ras.biodiversity.aq/aphia.php?p=taxdetails&id=238113 on 2026-09-09
Date
action
by
original description
Mrázek, A. (1893). Beitrag zur Kenntnis der Harpacticidenfauna des Süsswassers. [Contribution to the knowledge of the freshwater harpacticid fauna.]. <em>Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere.</em> 7(1): 89-130, pls. 4-7. (volume date published in 1894 however the first few issues were published in 20-v-1893)., available online at https://www.biodiversitylibrary.org/page/39199167
page(s): 108 [details]
basis of record Fiers, F. & V. Ghenne. (2000). Cryptozoic copepods from Belgium: diversity and biogeographic implications. <em>Belgian Journal of Zoology.</em> 103(1): 11-19. [details] Available for editors
additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors
additional source Bozkurt, A. (2017). First record of Epactophanes richardi Mrazek, 1893 (Copepoda, Harpacticoida, Camptocamptidae) for turkish inland waters. <em>Turkish Journal Of Fisheries And Aquatic Sciences.</em> 17(1):25-29., available online at https://doi.org/10.4194/1303-2712-v17_1_04 [details] Available for editors
additional source Brancelj, A., J. Lips & B. Lips. (2026). Cave Size Doesn't Matter but Persistence and Sampling Techniques Do—Rich Cave-Dwelling Fauna Revealed in the Epikarstic Velika Pasica Cave (Slovenia, Europe). <em>Diversity-Basel.</em> 18:1-21. May 2026., available online at https://doi.org/10.3390/d18050280 [details] Available for editors
additional source Brehm, V. (1912). Die Entomostraken der Danmark-Expedition. [The Entomostraca of the Denmark Expedition.]. <em>Meddelelser om Gronland, In: Danmark-Expeditionen til Gronlands Nordostkyst 1906-1908, Kobenhavn.</em> 5(5)45:305-317, pls.18-19. (1911). [details]
additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors
additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors
additional source Defaye, D. & B.H. Dussart. (2011). World Directory of Crustacea Copepoda III - Harpacticoida IV - Gelyelloida. <em>Backhuys Publishers, Margraf Publishers GmbH.</em> 450 pp.
note: "The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); ...
[details] Available for editors
additional source Dementev, M.S. (2015). Predvaritel'nyy obzor vodnykh bespozvonochnykh Tsentral'nogo Predkavkaz'ya (Severnyy Kavkaz) i prilegayushchikh gornykh territoriy. [Preliminary summary aquatic invertebrates in central Ciscaucasia (Nort Caucasus region) and adjacent mountain areas. ]. <em>Advances in Current Natural Sciences.</em> 6:102-118. [In Russian; English abstract]. [details] Available for editors
additional source Dussart, B.H. (1967). Les Copépodes des eaux continentales d'Europe occidentale. Tome I: Calanoïdes et Harpacticoïdes. [Copepods of the continental waters of Western Europe. Volume I: Calanoids and Harpacticoids.]. <em>N. Boubée et Cie, Paris.</em> 500 pp.
page(s): 290 [details] Available for editors
additional source Eldredge, L.G. & S.E. Miller. (1997). Numbers of Hawaiian species: Supplement 2, including a review of freshwater invertebrates. Occasional Papers of the Bernice P. Bishop Museum. Records of the Hawaii Biological Survey for 1996. Part I. Articles 48:3-22. [details] Available for editors
additional source Fefilova, E.B. & V.R. Alekseev. (2018). A new species and new records of harpacticoids (Crustacea: Copepoda: Harpacticoida) from North-Eastern Borneo. <em>Zoosystematica Rossica.</em> 27(2):205-217., available online at https://doi.org/10.31610/zsr/2018.27.2.205 [details] Available for editors
additional source Franke, U. (1989). Katalog zur Sammlung limnischer Copepoden von Prof. Dr. Friedrich Kiefer (Mit einem Gesamtverzeichnis seiner Publikationen). [Catolog of the freshwater copepods of Dr. Friedrich Kiefer.]. <em>Carolinea Beiheft, Karlsrühe.</em> 5:1-433, figs. 1-2.
note: Catalog vial number(s): 3532, 77 [details] Available for editors
additional source Ishida, T. & Y. Kikuchi. (2000). Illustrated fauna of the freshwater harpacticoid copepods of Japan. <em>Bulletin of the Biogeographical Society of Japan.</em> 55:7-94. (4.xii.2000). [details] Available for editors
additional source Ishida, T. (1982). Further notes on the freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 36:49-61, figs. 1-9. (iii-1982, Japanese with English summary.) [details] Available for editors
additional source Ishida, T. (1987). Freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 41:77-119. (Japanese summary.) [details] Available for editors
additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors
additional source Ishida, T. (1998). Freshwater copepods from the east coast of the Kamchatka Peninsula, Russia. In: Dahms, H.U., T. Glatzel, H.J. Hirche, S. Schiel & H.K. Schminke, (eds.). Proceedings of the 6th International Conference on Copepoda, Journal of Marine Systems, Spec. Vol. 15(1-4):391-396., available online at https://doi.org/10.1016/s0924-7963(97)00083-3 [details] Available for editors
additional source Karpowicz, M. & S. Watiroyram. (2026). Freshwater Harpacticoida from Borneo (Malaysia): New Records and Faunistic Insights into an Overlooked Group. <em>Diversity - Basel.</em> 18:1-13. Aug 2026., available online at https://doi.org/10.3390/d18080484 [details] Available for editors
additional source Kiefer, F. (1924). 3. Beiträge zur Copepodenkunde. 1. Canthocamptus pygmaeus Sars. 2. Ein neuer Fundort von Diaptomus denticornis Wierz. im Schwartzwald. 3. Epactophanes richardi Mràzek. [3. Contributions to the study of copepods. 1. Canthocamptus pygmaeus Sars. 2. A new locality for Diaptomus denticornis Wierz. in the Black Forest. 3. Epactophanes richardi Mràzek.]. <em>Zoologischer Anzeiger.</em> 59(7-8):200-204, figs. 1-8. (May1924). [details] Available for editors
additional source Kiefer, F. (1960). Psammobionte Ruderfusskrebse (Crust. Cop.) aus dem Gebiet der Unterweser und von der Insel Helgoland. [Psammobiont copepods (Crust. Cop.) from the Lower Weser region and from the island of Heligoland. <em>Zoologischer Anzeiger.</em> 165(1-2):30-36, figs. 1-39. (vii-1960). [details] Available for editors
additional source Kiefer, F. (1963). Über einige Copepoda Harpacticoida aus grabungen an norditalienischen Fliessgewässern. [On some Copepoda Harpacticoida from excavations at northern Italian rivers.]. <em>Estratto dalle Memorie del Museo Civico di Storia Naturale, Verona.</em> 11:1-10. (15 Jul 1963). [details] Available for editors
additional source Kulhavy, V. (1957). Plazivky (Copepoda-Harpacticoidea) jihoceskych mechu. [Harpacticoid copepods from South Bohemian mosses.]. <em>Acta Musei Nationalis, Prahba, [ Casopis Národniho Musea].</em> 126(1):35-43, figs. 1-6. (Czech with English summary.).
page(s): 41 [details] Available for editors
additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors
additional source Lewis, M.H. (1980). Systematic studies on the New Zealand freshwater Harpacticoida (Crustacea: Copepoda). <em>Ph.D. Thesis, University of Auckland, Auckland, New Zealand.</em> 314 pp., available online at https://hdl.handle.net/2292/1599 [details]
additional source Marin, I., D. Sharafutdinova, T. Mukhina, A. Chernyshev & A. Novikov. (2022). The record of Epactophanes richardi Mrázek, 1893 (Harpacticoida: Canthocamptidae) in soil of the Russian Far-Eastern forest. <em>Journal of Asia-Pacific Biodiversity.</em> 15:454-458., available online at https://doi.org/10.1016/j.japb.2022.05.002 [details] Available for editors
additional source Menzel, R. (1914). Zur Kenntnis von Moraria muscicola Richters nebst Hinweis auf deren Verwandtschaft mit Epactophanes richardi Mrázek. [On the knowledge of Moraria muscicola Richters including reference to its relationship with Epactophanes richardi Mrázek.]. <em>Zoologischer Anzeiger.</em> 43(13):615-620, figs. 1-5. (17-iii-1914). [details]
additional source Michailova-Neikova, M. (1963). Epactophanes richardi Mrázek. <em>Priroda, Moscow.</em> 12(4):101-102. [details]
additional source Monchenko, V.I. (1964). Novi dlya Ukraini veslonogi rakopodibni (Copepoda): Parastenocaris brevipes ta Epactophanes richardi. [New copepods for the Ukraine: Parastenocaris brevipes and Epactophanes richardi.]. <em>Dopovidi Akademiyi Nauk Ukrayins'koiy RSR, Kiev.</em> 1964(4):545-549. (Ukrainian with Russian and English summaries). [details] Available for editors
additional source Okuneva, G.L. (1989). Garpaktitsidy ozera Bakala. [The harpacticoids of Lake Baikal.]. <em>Izdatel'stvo Instituta, Irkutsk.</em> 150 pp. (In Russian.). [details] Available for editors
additional source Reid, B.L. (2007). Energy flow in a floodplain aquifer ecosystem. <em>Ph.D. Dissertation, University of Montana, Missoula, Montana, U.S.A.</em> 250 pp., available online at https://scholarworks.umt.edu/etd/372 [details]
additional source Scher, O., D. Defaye, N.M. Korovchinsky & A. Thiéry. (2000). The crustacean fauna (Branchiopoda, Copepoda) of shallow freshwater bodies in Iceland. Vestnik Zoologii, Kiev 34(6):11-25. (7.xi.2000) [details] Available for editors
additional source Willems, K.A. (1989). Verspreiding, ecologie en gemeenschapsstructuur van benthische copepoden in het Delta-gebied en de Eems-Dollard (Nederland). [Distribution, ecology and community structure of benthic copepods in the Delta area and the Ems-Dollard (Netherlands)]. <em>Ph.D. Thesis, Universiteit Gent, Ghent, Belgium.</em> 2 vols:198 pp., available online at http://www.vliz.be/en/imis?module=dataset&dasid=5080) [details] Available for editors
page(s): 108 [details]
basis of record Fiers, F. & V. Ghenne. (2000). Cryptozoic copepods from Belgium: diversity and biogeographic implications. <em>Belgian Journal of Zoology.</em> 103(1): 11-19. [details] Available for editors
additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors
additional source Bozkurt, A. (2017). First record of Epactophanes richardi Mrazek, 1893 (Copepoda, Harpacticoida, Camptocamptidae) for turkish inland waters. <em>Turkish Journal Of Fisheries And Aquatic Sciences.</em> 17(1):25-29., available online at https://doi.org/10.4194/1303-2712-v17_1_04 [details] Available for editors
additional source Brancelj, A., J. Lips & B. Lips. (2026). Cave Size Doesn't Matter but Persistence and Sampling Techniques Do—Rich Cave-Dwelling Fauna Revealed in the Epikarstic Velika Pasica Cave (Slovenia, Europe). <em>Diversity-Basel.</em> 18:1-21. May 2026., available online at https://doi.org/10.3390/d18050280 [details] Available for editors
additional source Brehm, V. (1912). Die Entomostraken der Danmark-Expedition. [The Entomostraca of the Denmark Expedition.]. <em>Meddelelser om Gronland, In: Danmark-Expeditionen til Gronlands Nordostkyst 1906-1908, Kobenhavn.</em> 5(5)45:305-317, pls.18-19. (1911). [details]
additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors
additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors
additional source Defaye, D. & B.H. Dussart. (2011). World Directory of Crustacea Copepoda III - Harpacticoida IV - Gelyelloida. <em>Backhuys Publishers, Margraf Publishers GmbH.</em> 450 pp.
note: "The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); ...
"The type species of this genus is Epactophannes richardi Mrázek, 1893. Numerous varieties and subspecies of this unique representative of the genus have been described. Lang (1948), Dussart (1967); Damian-Georgescu (1970) and Tai & Song (1979) considered them as synonyms of the nominal species which appears to have an important morphological variability. Borutzky (1952) did not agree with this opinion" (Defaye & Dussart, 2011: 115).
additional source Dementev, M.S. (2015). Predvaritel'nyy obzor vodnykh bespozvonochnykh Tsentral'nogo Predkavkaz'ya (Severnyy Kavkaz) i prilegayushchikh gornykh territoriy. [Preliminary summary aquatic invertebrates in central Ciscaucasia (Nort Caucasus region) and adjacent mountain areas. ]. <em>Advances in Current Natural Sciences.</em> 6:102-118. [In Russian; English abstract]. [details] Available for editors
additional source Dussart, B.H. (1967). Les Copépodes des eaux continentales d'Europe occidentale. Tome I: Calanoïdes et Harpacticoïdes. [Copepods of the continental waters of Western Europe. Volume I: Calanoids and Harpacticoids.]. <em>N. Boubée et Cie, Paris.</em> 500 pp.
page(s): 290 [details] Available for editors
additional source Eldredge, L.G. & S.E. Miller. (1997). Numbers of Hawaiian species: Supplement 2, including a review of freshwater invertebrates. Occasional Papers of the Bernice P. Bishop Museum. Records of the Hawaii Biological Survey for 1996. Part I. Articles 48:3-22. [details] Available for editors
additional source Fefilova, E.B. & V.R. Alekseev. (2018). A new species and new records of harpacticoids (Crustacea: Copepoda: Harpacticoida) from North-Eastern Borneo. <em>Zoosystematica Rossica.</em> 27(2):205-217., available online at https://doi.org/10.31610/zsr/2018.27.2.205 [details] Available for editors
additional source Franke, U. (1989). Katalog zur Sammlung limnischer Copepoden von Prof. Dr. Friedrich Kiefer (Mit einem Gesamtverzeichnis seiner Publikationen). [Catolog of the freshwater copepods of Dr. Friedrich Kiefer.]. <em>Carolinea Beiheft, Karlsrühe.</em> 5:1-433, figs. 1-2.
note: Catalog vial number(s): 3532, 77 [details] Available for editors
additional source Ishida, T. & Y. Kikuchi. (2000). Illustrated fauna of the freshwater harpacticoid copepods of Japan. <em>Bulletin of the Biogeographical Society of Japan.</em> 55:7-94. (4.xii.2000). [details] Available for editors
additional source Ishida, T. (1982). Further notes on the freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 36:49-61, figs. 1-9. (iii-1982, Japanese with English summary.) [details] Available for editors
additional source Ishida, T. (1987). Freshwater harpacticoid copepods of Hokkaido, northern Japan. Scientific Reports of the Hokkaido Salmon Hatchery 41:77-119. (Japanese summary.) [details] Available for editors
additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors
additional source Ishida, T. (1998). Freshwater copepods from the east coast of the Kamchatka Peninsula, Russia. In: Dahms, H.U., T. Glatzel, H.J. Hirche, S. Schiel & H.K. Schminke, (eds.). Proceedings of the 6th International Conference on Copepoda, Journal of Marine Systems, Spec. Vol. 15(1-4):391-396., available online at https://doi.org/10.1016/s0924-7963(97)00083-3 [details] Available for editors
additional source Karpowicz, M. & S. Watiroyram. (2026). Freshwater Harpacticoida from Borneo (Malaysia): New Records and Faunistic Insights into an Overlooked Group. <em>Diversity - Basel.</em> 18:1-13. Aug 2026., available online at https://doi.org/10.3390/d18080484 [details] Available for editors
additional source Kiefer, F. (1924). 3. Beiträge zur Copepodenkunde. 1. Canthocamptus pygmaeus Sars. 2. Ein neuer Fundort von Diaptomus denticornis Wierz. im Schwartzwald. 3. Epactophanes richardi Mràzek. [3. Contributions to the study of copepods. 1. Canthocamptus pygmaeus Sars. 2. A new locality for Diaptomus denticornis Wierz. in the Black Forest. 3. Epactophanes richardi Mràzek.]. <em>Zoologischer Anzeiger.</em> 59(7-8):200-204, figs. 1-8. (May1924). [details] Available for editors
additional source Kiefer, F. (1960). Psammobionte Ruderfusskrebse (Crust. Cop.) aus dem Gebiet der Unterweser und von der Insel Helgoland. [Psammobiont copepods (Crust. Cop.) from the Lower Weser region and from the island of Heligoland. <em>Zoologischer Anzeiger.</em> 165(1-2):30-36, figs. 1-39. (vii-1960). [details] Available for editors
additional source Kiefer, F. (1963). Über einige Copepoda Harpacticoida aus grabungen an norditalienischen Fliessgewässern. [On some Copepoda Harpacticoida from excavations at northern Italian rivers.]. <em>Estratto dalle Memorie del Museo Civico di Storia Naturale, Verona.</em> 11:1-10. (15 Jul 1963). [details] Available for editors
additional source Kulhavy, V. (1957). Plazivky (Copepoda-Harpacticoidea) jihoceskych mechu. [Harpacticoid copepods from South Bohemian mosses.]. <em>Acta Musei Nationalis, Prahba, [ Casopis Národniho Musea].</em> 126(1):35-43, figs. 1-6. (Czech with English summary.).
page(s): 41 [details] Available for editors
additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors
additional source Lewis, M.H. (1980). Systematic studies on the New Zealand freshwater Harpacticoida (Crustacea: Copepoda). <em>Ph.D. Thesis, University of Auckland, Auckland, New Zealand.</em> 314 pp., available online at https://hdl.handle.net/2292/1599 [details]
additional source Marin, I., D. Sharafutdinova, T. Mukhina, A. Chernyshev & A. Novikov. (2022). The record of Epactophanes richardi Mrázek, 1893 (Harpacticoida: Canthocamptidae) in soil of the Russian Far-Eastern forest. <em>Journal of Asia-Pacific Biodiversity.</em> 15:454-458., available online at https://doi.org/10.1016/j.japb.2022.05.002 [details] Available for editors
additional source Menzel, R. (1914). Zur Kenntnis von Moraria muscicola Richters nebst Hinweis auf deren Verwandtschaft mit Epactophanes richardi Mrázek. [On the knowledge of Moraria muscicola Richters including reference to its relationship with Epactophanes richardi Mrázek.]. <em>Zoologischer Anzeiger.</em> 43(13):615-620, figs. 1-5. (17-iii-1914). [details]
additional source Michailova-Neikova, M. (1963). Epactophanes richardi Mrázek. <em>Priroda, Moscow.</em> 12(4):101-102. [details]
additional source Monchenko, V.I. (1964). Novi dlya Ukraini veslonogi rakopodibni (Copepoda): Parastenocaris brevipes ta Epactophanes richardi. [New copepods for the Ukraine: Parastenocaris brevipes and Epactophanes richardi.]. <em>Dopovidi Akademiyi Nauk Ukrayins'koiy RSR, Kiev.</em> 1964(4):545-549. (Ukrainian with Russian and English summaries). [details] Available for editors
additional source Okuneva, G.L. (1989). Garpaktitsidy ozera Bakala. [The harpacticoids of Lake Baikal.]. <em>Izdatel'stvo Instituta, Irkutsk.</em> 150 pp. (In Russian.). [details] Available for editors
additional source Reid, B.L. (2007). Energy flow in a floodplain aquifer ecosystem. <em>Ph.D. Dissertation, University of Montana, Missoula, Montana, U.S.A.</em> 250 pp., available online at https://scholarworks.umt.edu/etd/372 [details]
additional source Scher, O., D. Defaye, N.M. Korovchinsky & A. Thiéry. (2000). The crustacean fauna (Branchiopoda, Copepoda) of shallow freshwater bodies in Iceland. Vestnik Zoologii, Kiev 34(6):11-25. (7.xi.2000) [details] Available for editors
additional source Willems, K.A. (1989). Verspreiding, ecologie en gemeenschapsstructuur van benthische copepoden in het Delta-gebied en de Eems-Dollard (Nederland). [Distribution, ecology and community structure of benthic copepods in the Delta area and the Ems-Dollard (Netherlands)]. <em>Ph.D. Thesis, Universiteit Gent, Ghent, Belgium.</em> 2 vols:198 pp., available online at http://www.vliz.be/en/imis?module=dataset&dasid=5080) [details] Available for editors
Present Inaccurate Introduced: alien Containing type locality
